J. For. Sci., 2026, 72(2):73-81 | DOI: 10.17221/92/2025-JFS

Altitudinal patterns of woody vegetation diversity in Al-Jabal Al-Akhdar, a Mediterranean mountainOriginal Paper

Moussa Masoud ORCID...1,2, Hazandy Abdul-Hamid ORCID...1,3, Johar Mohamed ORCID...1,3, Attia Alsanousi ORCID...1,2
1 Department of Forestry Science and Biodiversity, Faculty of Forestry and Environment, Universiti Putra Malaysia, Serdang, Selangor, Malaysia
2 Department of Forestry, Faculty of Natural Resources and Environmental Sciences, University of Omar Al-Mukhtar Al-Bayda, Bayda, Libya
3 Laboratory of Bioresource Management, Institute of Tropical Forestry and Forest Products, Universiti Putra Malaysia, Serdang, Selangor, Malaysia

This study examines the floristic composition, structural attributes, diversity patterns, and distribution of endemic woody species along an altitudinal gradient in the Al-Jabal Al-Akhdar mountain of northeastern Libya. Stratified sampling was conducted across four elevation zones on the northern and southern slopes. Phytosociological parameters – including density, basal area, importance value index, and diversity indices – were assessed, and Pearson correlations were used to evaluate relationships with altitude and aspect. A total of 61 woody species from 43 genera and 26 families were recorded, highlighting the region's biodiversity. Tree and shrub density and basal area increased with elevation, particularly on northern slopes. Juniperus phoenicea L. emerged as the most dominant tree, while Phlomis floccosa D. Don and Pistacia lentiscus L. dominated the shrub layer. Diversity indices showed negative correlations with altitude, indicating declining species richness and evenness at higher elevations under cooler temperatures and greater environmental stress. Ten endemic woody species were documented, with endemic richness positively associated with altitude, reflecting the role of habitat isolation and environmental filtering. These findings provide essential insights for biodiversity conservation, climate adaptation, and ecosystem management in Al-Jabal Al-Akhdar and comparable mountainous ecosystems.

Keywords: elevation; forest ecology; phytosociology; species richness

Received: December 7, 2025; Revised: January 30, 2026; Accepted: February 2, 2026; Prepublished online: February 26, 2026; Published: February 27, 2026  Show citation

ACS AIP APA ASA Harvard Chicago Chicago Notes IEEE ISO690 MLA NLM Turabian Vancouver
Masoud M, Abdul-Hamid H, Mohamed J, Alsanousi A. Altitudinal patterns of woody vegetation diversity in Al-Jabal Al-Akhdar, a Mediterranean mountain. Journal of Forest Science. 2026;72(2):73-81. doi: 10.17221/92/2025-JFS.
Download citation

References

  1. Al-Jabal Al-Akhdar South Project (2005): Study and Evaluation of Natural Vegetation in the Al-Jabal Al-Akhdar Area. Al Bayda, Omar Al-Mokhtar University: 946. (in Arabic)
  2. Alsanousi A.A., Abdul-Hamid H., Mohamed J., Masoud M. (2025): Pinus halepensis Mill. in the Mediterranean region: A review of ecological significance, growth patterns, and soil interactions. iForest, 18: 30-37. Go to original source...
  3. Boulos L. (1972): Our present knowledge on the flora and vegetation of Libya bibliography. Webbia, 26: 365-400. Go to original source...
  4. Boulos L. (1997): Endemic flora of the Middle East and North Africa. In: Barakat H.N., Hegazy A.K. (eds): Reviews in Ecology: Desert Conservation and Development. Cairo, Metropole: 229-260.
  5. Braun K., Passon J., Jeworutzki A. (2020): Across the vast land - Some aspects on Libya's geography. In: Braun K., Passon J. (eds): Across the Sahara. Cham, Springer: 1-28. Go to original source...
  6. Jafri S., El-Gadi A. (1993): Flora of Libya. Tripoli, Al Fatteh University, Faculty of Science, Botany Department: 4625.
  7. Malik Z.A., Bhat J.A., Ballabha R., Bussmann R.W., Bhatt A.B. (2015): Ethnomedicinal plants traditionally used in health care practices by inhabitants of Western Himalaya. Journal of Ethnopharmacology, 172: 133-144. Go to original source... Go to PubMed...
  8. Margalef D.R. (1958): Information theory in ecology. General Systems, 3: 36e71.
  9. Masoud M. (2012): Influence of climatic zones on the distribution and abundance of damage agents and forest types in Colorado, United States and Jalisco, Mexico. [MSc. Thesis] Fort Collins, Colorado State University.
  10. Masoud M. (2016): Monitoring land use/land cover using multi-temporal Landsat images in Al-Jabal Al-Akhdar area in Libya between 1984 and 2003. Al-Mukhtar Journal of Sciences, 31: 12-23. Go to original source...
  11. Masoud M., Abdul-Hamid H., Bin Mohamed J., Alsanousi A. (2024): Investigating soil properties on the north and south slopes at different elevations in Al-Jabal Al-Akhdar, Libya. Forest Science and Technology, 20: 286-299. Go to original source...
  12. Masoud M., Abdul-Hamid H., Mohamed J., Alsanousi A. (2025): Altitude-induced variations in vegetation characteristics and soil properties. Nature Environment and Pollution Technology, 24: D1773. Go to original source...
  13. Phillips E.A. (1959): Methods of Vegetation Study. New York, Henry Holt and Co.: 107.
  14. Pielou E.C. (1975): Ecological Diversity. New York, Wiley: 165.
  15. R Core Team (2024): R: A language and environment for statistical computing. Vienna, R Foundation for Statistical Computing. Available at: http://www.R-project.org
  16. Shannon C.E., Wiener W. (1963): The Mathematical Theory of Communication. Urbana, University of Illinois Press: 127.
  17. Simpson E.H. (1949): Measurement of diversity. Nature, 163: 688e690. Go to original source...
  18. Sinha S., Badola H.K., Chhetri B., Gaira K.S., Lepcha J., Dhyani P.P. (2018): Effect of altitude and climate in shaping the forest compositions of Singalila National Park in Khangchendzonga landscape, Eastern Himalaya, India. Journal of Asia-Pacific Biodiversity, 11: 267-275. Go to original source...
  19. Steinbauer M.J., Field R., Grytnes J.A., Trigas P., Ah-Peng C., Attorre F., Birks H.J.B., Borges P.A., Cardoso P., Chou C.H., Beierkuhnlein C. (2016): Topography-driven isolation, speciation and a global increase of endemism with elevation. Global Ecology and Biogeography, 25: 1097-1107. Go to original source...
  20. Theurillat J.P., Guisan A. (2001): Potential impact of climate change on vegetation in the European Alps: A review. Climatic Change, 50: 77-109. Go to original source...
  21. Vetaas O.R., Grytnes J.A. (2002): Distribution of vascular plant species richness and endemic richness along the Himalayan elevation gradient in Nepal. Global Ecology and Biogeography, 11: 291-301. Go to original source...

This is an open access article distributed under the terms of the Attribution-NonCommercial 4.0 International (CC BY-NC 4.0.), which permits non-comercial use, distribution, and reproduction in any medium, provided the original publication is properly cited. No use, distribution or reproduction is permitted which does not comply with these terms.