J. For. Sci., 2023, 69(11):470-484 | DOI: 10.17221/76/2023-JFS

Potential of mixed Picea abies (L.) Karst. and Pinus sylvestris L. forests in lowland areas of Central BohemiaOriginal Paper

Pavel Brabec, Jakub Brichta, Zdeněk Vacek, Stanislav Vacek, Václav Šimůnek, Vojtěch Hájek
Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Prague, Czech Republic

Mixed forests play a key role in terms of stability, production potential, and adaptation to climate change. In addition, the studied Norway spruce [Picea abies (L.) Karst.] and Scots pine (Pinus sylvestris L.) belong to the most economically important tree species in Europe. The objectives were to determine the effect of the species composition of these two tree genera on the production, structure, diversity, and growth of mixed stands at lower elevations in the Czech Republic. Based on dendroecological samples, research was also carried out on the influence of climatic factors and climate change on the radial growth of these trees of interest. Mixed forests showed higher timber production by 29.8% compared to spruce and pine monocultures. The production of mature stands ranged from 328 to 479 m3·ha–1. Spruce achieved higher radial growth, but its growth variability was higher than that of pine. Compared to precipitation, temperatures had a greater influence on the radial growth of both tree species, especially in the growing season. In terms of diversity, mixed stands achieved significantly higher structural differentiation and overall diversity compared to monospecific variants. Mixed stands can achieve higher production potential, diversity, and especially resistance to climatic extremes in the lowland regions of the Czech Republic. The differences between mixed stands and monocultures, i.e. the effect of tree species mixing, depend on the appropriate proportions of tree species and their spatial pattern.

Keywords: climate change; dendrochronology; Norway spruce; productivity; Scots pine

Received: June 30, 2023; Revised: August 9, 2023; Accepted: August 18, 2023; Prepublished online: November 13, 2023; Published: November 27, 2023  Show citation

ACS AIP APA ASA Harvard Chicago Chicago Notes IEEE ISO690 MLA NLM Turabian Vancouver
Brabec P, Brichta J, Vacek Z, Vacek S, Šimůnek V, Hájek V. Potential of mixed Picea abies (L.) Karst. and Pinus sylvestris L. forests in lowland areas of Central Bohemia. J. For. Sci. 2023;69(11):470-484. doi: 10.17221/76/2023-JFS.
Download citation

References

  1. Abdullah H., Skidmore A.K., Darvishzadeh R., Heurich M. (2019): Sentinel-2 accurately maps green attack stage of European spruce bark beetle (Ips typographus L.) compared with Landsat-8. Remote Sensing in Ecology and Conservation, 5: 87-106. Go to original source...
  2. Albrich K., Rammer W., Seidl R. (2020): Climate change causes critical transitions and irreversible alterations of mountain forests. Global Change Biology, 26: 4013-4027. Go to original source... Go to PubMed...
  3. Aleksandrowicz-Trzcińska M., Drozdowski S., Brzeziecki B., Rutkowska P., Jabłońska B. (2014): Effects of different methods of site preparation on natural regeneration of Pinus sylvestris in Eastern Poland. Dendrobiology, 71: 73-81. Go to original source...
  4. Aleksandrowicz-Trzcińska M., Drozdowski S., Wołczyk Z., Bielak K., Żybura H. (2017): Effects of reforestation and site preparation methods on early growth and survival of Scots pine (Pinus sylvestris L.) in south-eastern Poland. Forests, 8: 1-17. Go to original source...
  5. Aleksandrowicz-Trzcińska M., Drozdowski S., Studnicki M., Żybura H. (2018): Effects of site preparation methods on the establishment and natural-regeneration traits of Scots pines (Pinus sylvestris L.) in northeastern Poland. Forests, 9: 717. Go to original source...
  6. Allen C.D., Breshears D.D., McDowell N.G. (2015): On underestimation of global vulnerability to tree mortality and forest die-off from hotter drought in the Anthropocene. Ecosphere, 6: 1-55. Go to original source...
  7. Andrzejczyk T. (2006): Rębnia przerębowa w drzewostanach sosnowych. Sylwan, 150: 52-60. (in Polish)
  8. Augustaitis A., Augustaitiene I., Deltuvas R. (2007): Scots pine (Pinus sylvestris L.) crown defoliation in relation to the acid deposition and meteorology in Lithuania. Water, Air, and Soil Pollution, 182: 335-348. Go to original source...
  9. Barta M., Lalik M., Rell S., Kunca A., Horáková M.K., Mudrončenková S., Galko J. (2019): Hypocrealean fungi associated with Hylobius abietis in Slovakia, their virulence against weevil adults and effect on feeding damage in laboratory. Forests, 10: 634. Go to original source...
  10. Bauhus J., Forrester D.I., Gardiner B., Jactel H., Vallejo R., Pretzsch H. (2017): Ecological stability of mixed-species forests. In: Pretzsch H., Forrester D., Bauhus J. (eds): Mixed-Species Forests: Ecology and Management. Berlin, Springer: 337-382. Go to original source...
  11. Bielak K., Dudzińska M., Pretzsch H. (2014): Mixed stands of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies [L.] Karst.) can be more productive than monocultures. Evidence from over 100 years of observation of long-term experiments. Forest Systems, 23: 573-589. Go to original source...
  12. Bílek L., Remeš J., Švec O., Zahradník D. (2013): On the way to continuous cover forest at middle elevations - The question of forest structure and specific site characteristics. Journal of Forest Science, 59: 391-397. Go to original source...
  13. Bílek L., Vacek S., Vacek Z., Remeš J., Král J., Bulušek D., Gallo J. (2016): How close to nature is close-to-nature pine silviculture? Journal of Forest Science, 62: 24-34. Go to original source...
  14. Biondi F., Waikul K. (2004): Dendroclim 2002: AC++ program for statistical calibration of climate signals in tree ring chronologies. Computers and Geosciences, 30: 303-311. Go to original source...
  15. Bogino S., Fernández Nieto M.J., Bravo F. (2009): Climate effect on radial growth of Pinus sylvestris at its southern and western distribution limits. Silva Fennica, 43: 609-623. Go to original source...
  16. Brabec P., Vacek Z., Vacek S., Štefančík I., Cukor J., Weatherall A., Gallo J., Slávik M., Sitková Z., Putalová T. (2023): Growth-climate responses of Picea sitchensis (Bong.) Carr. versus Picea abies (L.) Karst. in the British Isles and Central Europe. Central European Forestry Journal, 69: 167-178. Go to original source...
  17. Brichta J., Bílek L., Linda R., Vítámvás J. (2020): Does shelterwood regeneration on natural Scots pine sites under changing environmental conditions represent a viable alternative to traditional clear-cut management? Central European Forestry Journal, 66: 104-115. Go to original source...
  18. Brichta J., Vacek S., Vacek Z., Cukor J., Mikeska M., Bílek L., Šimůnek V., Gallo J., Brabec P. (2023): Importance and potential of Scots pine (Pinus sylvestris L.) in 21st century. European Forestry Journal, 69: 3-20. Go to original source...
  19. Bublinec E. (1994): Koncentrácia, akumulácia a kolobeh prvkov v bukovom a smrekovom ekosystéme. Bratislava, VEDA: 85. (in Slovak)
  20. Bunn A., Mikko K. (2018): Chronology Building in dplR. Vienna, R Foundation for Statistical Computing: 13.
  21. Buras A., Schunk C., Zeiträg C., Herrmann C., Kaiser L., Lemme H., Straub C., Taeger S., Gößwein S., Klemmt H., Menzel A. (2018): Are Scots pine forest edges particularly prone to drought-induced mortality? Environmental Research Letters, 13: 025001. Go to original source...
  22. Camarero J.J., Gazol A., Sangüesa-Barreda G., Cantero A., Sánchez-Salguero R., Sánchez-Miranda A., Granda E., Serra-Maluquer X., Ibáñez R. (2018): Forest growth responses to drought at short- and long-term scales in Spain: Squeezing the stress memory from tree rings. Frontiers in Ecology and Evolution, 6: 9. Go to original source...
  23. Caudullo G., Tinner W., de Rigo D. (2016): Picea abies in Europe: Distribution, habitat, usage and threats. In: San-Miguel-Ayanz J., de Rigo D., Caudullo G., Houston Durrant T., Mauri A. (eds): European Atlas of Forest Tree Species. Luxembourg, Publications Office of the European Union: 114-116.
  24. Čermák P., Kolář T., Žid T., Trnka M., Rybníček M. (2019): Norway spruce responses to drought forcing in areas affected by forest decline. Forest Systems, 28: e016. Go to original source...
  25. Čihák T., Vejpustková M. (2021): Produkční charakteristiky borovice lesní v České republice na základě informací z druhého cyklu Národní inventarizace lesů (2011-2015). Zprávy lesnického výzkumu, 66: 126-137. (in Czech)
  26. Čížek J., Kratochvíl F., Peřina V. (1959): Přeměny monokultur. Praha, SZN: 191. (in Czech)
  27. Clark P.J., Evans F.C. (1954): Distance to nearest neighbor as a measure of spatial relationships in populations. Ecology, 35: 445-453. Go to original source...
  28. Cook E.R., Holmes R.L. (1984): Program ARSTAN user manual. Tucson, Laboratory of Tree-Ring Research, University of Arizona: 15.
  29. Crookston N.L., Stage A.R. (1999): Percent Canopy Cover and Stand Structure Statistics from the Forest Vegetation Simulator. Ogden, Rocky Mountain Research Station: 11. Go to original source...
  30. Cukor J., Vacek Z., Linda R., Vacek S., Marada P., Šimůnek V., Havránek F. (2019a): Effects of bark stripping on timber production and structure of Norway spruce forests in relation to climatic factors. Forests, 10: 320. Go to original source...
  31. Cukor J., Vacek Z., Linda R., Sharma R.P., Vacek S. (2019b): Afforested farmland vs. forestland: Effects of bark stripping by Cervus elaphus and climate on production potential and structure of Picea abies forests. PLoS ONE, 14: e0221082. Go to original source... Go to PubMed...
  32. Cukor J., Vacek Z., Linda R., Vacek S., Šimůnek V., Macháček Z., Brichta J., Prokůpková A. (2022): Scots pine (Pinus sylvestris L.) demonstrates a high resistance against bark stripping damage. Forest Ecology and Management, 513: 120182. Go to original source...
  33. D'Andrea G., Šimůnek V., Pericolo O., Vacek Z., Vacek S., Corleto R., Olejár L., Ripullone F. (2023): Growth response of Norway spruce (Picea abies [L.] Karst.) in central Bohemia (Czech Republic) to climate change. Forests, 14: 1215. Go to original source...
  34. De Vries W., Dobbertin M.H., Solberg S., Van Dobben H.F., Schaub M. (2014): Impacts of acid deposition, ozone exposure and weather conditions on forest ecosystems in Europe: An overview. Plant and Soil, 380: 1-45. Go to original source...
  35. del Río M., Pretzsch H., Alberdi I., Bielak K., Bravo F., Brunner A., Condés S., Ducey M.J., Fonseca T., von Lüpke N. et al. (2016): Characterization of the structure, dynamics, and productivity of mixed-species stands: Review and perspectives. European Journal of Forest Research, 135: 23-49. Go to original source...
  36. Desplanque C., Rolland C., Schweingruber F.H. (1999): Influence of species and abiotic factors on extreme tree ring modulation. Trees., 13: 218-227. Go to original source...
  37. Drexhage M., Colin F. (2001): Estimating root system biomass from breast-height diameters. Forestry, 74: 491-497. Go to original source...
  38. Drössler L., Agestam E., Bielak K., Dudzinska M., Koricheva J., Liziniewicz M., Löf M., Mason B., Pretzsch H., Valkonen S., Wellhausen K. (2018): Over-and underyielding in time and space in experiments with mixed stands of Scots pine and Norway spruce. Forests, 9: 495. Go to original source...
  39. Durant D.T., de Rigo D., Caudullo G. (2016): Pinus sylvestris in Europe: Distribution, habitat, usage and threats. In: San-Miguel-Ayanz J., de Rigo D., Caudullo G., Houston Durrant T., Mauri A. (eds): European Atlas of Forest Tree Species. Luxembourg, Publications Office of the European Union: 114-116.
  40. Dyderski M.K., Paz S., Frelich L.E., Jagodziński A.M. (2018): How much does climate change threaten European forest tree species distributions? Global Change Biology, 24: 1150-1163. Go to original source... Go to PubMed...
  41. Eckenwalder J.E. (2009): Conifers of the World: The Complete Reference. Portland, Timber Press: 720.
  42. Fabrika M., Ďurský J. (2005): Algorithms and software solution of thinning models for SIBYLA growth simulator. Journal of Forensic Sciences, 51: 431-445. Go to original source...
  43. Farjon A. (2010): A Handbook of the World's Conifers. Leiden, Brill: 1111. Go to original source...
  44. Flexas J., Medrano H. (2002): Drought-inhibition of photosynthesis in C3 plants: Stomatal and non-stomatal limitations revisited. Annals of Botany, 89: 183-189. Go to original source... Go to PubMed...
  45. Fritts H.C. (1976): Tree Rings and Climate. London, Academic Press: 567.
  46. Füldner K. (1995): Strukturbeschreibung in Mischbeständen. Forstarchiv, 66: 235-606. (in German)
  47. Fürst C., Vacik H., Lorz C., Makeschin F., Podrázský V., Janeček V. (2007): Meeting the challenges of process-oriented forest management. Forest Ecology and Management, 248: 1-5. Go to original source...
  48. Gallo J., Bílek L., Šimůnek V., Roig S., Fernández J.A.B. (2020): Uneven-aged silviculture of Scots pine in Bohemia and Central Spain: Comparison study of stand reaction to transition and long-term selection management. Journal of Forest Science, 66: 22-35. Go to original source...
  49. Gallo J., Vacek Z., Vacek S. (2021): Quarter of a century of forest fertilization and liming research at the Department of Silviculture in Prague, Czech Republic. Central European Forestry Journal, 67: 123-134. Go to original source...
  50. Gao Y., Markkanen T., Aurela M., Mammarella I., Thum T., Tsuruta A., Yang H., Aalto T. (2017): Response of water use efficiency to summer drought in a boreal Scots pine forest in Finland. Biogeosciences, 14: 4409-4422. Go to original source...
  51. Gazol A., Camarero J.J., Anderegg W.R.L., Vicente-Serrano S.M. (2017): Impacts of droughts on the growth resilience of Northern Hemisphere forests. Global Ecology and Biogeography, 26: 166-176. Go to original source...
  52. Greenwood S., Ruiz-Benito P., Martínez-Vilalta J., Lioret F., Kitzberge T., Alle C.D., Fensham R., Laughlin D.C., Kattge J., Bönisch G., Kraft N.J.B., Jump A.S. (2017): Tree mortality across biomes is promoted by drought intensity, lower wood density and higher specific leaf area. Ecology Letters, 20: 539-553. Go to original source... Go to PubMed...
  53. Grissino-Mayer H.D., Holmes R.L., Fritts H.C. (1992): International Tree-Ring Data Bank Program Library: User's Manual. Tuscon, Laboratory of Tree-Ring Research, University of Arizona: 106.
  54. Haberstroh S., Werner C., Grün M., Kreuzwieser J., Seifert T., Schindler D., Christen A. (2022): Central European 2018 hot drought shifts Scots pine forest to its tipping point. Plant Biology, 7: 1186-1197. Go to original source... Go to PubMed...
  55. Halaj J., Grék J., Pánek F., Petráš R., Řehák J. (1987): Rastové tabuľky hlavných drevín ČSSR. Bratislava, Príroda: 361. (in Slovak)
  56. Hanewinkel M., Pretzsch H. (2000): Modelling the conversion from even-aged to uneven-aged stands of Norway spruce (Picea abies L. KARST.) with a distance-dependent growth simulator. Forest Ecology and Management, 134: 55-70. Go to original source...
  57. Hanewinkel M., Cullmann D.A., Schelhaas M.J., Nabuurs G.J., Zimmermann N.E. (2013): Climate change may cause severe loss in the economic value of European forest land. Nature Climate Change, 3: 203-207. Go to original source...
  58. Hartl-Meier C., Zang C., Büntgen U., Esper J., Rothe A., Göttlein A., Dirnböck T., Treydte K. (2014): Uniform climate sensitivity in tree-ring stable isotopes across species and sites in a mid-latitude temperate forest. Tree Physiology, 35: 4-15. Go to original source... Go to PubMed...
  59. Hlásny T., Krokene P., Liebhold A., Montagné-Huck C., Müller J., Qin H., Raffa K., Schelhaas M.-J., Seidl R., Svoboda M., Viiri H. (2019): Living with Bark Beetles: Impacts, Outlook and Management Options. From Science to Policy 8. Joensuu, European Forest Institute: 50. Go to original source...
  60. Hlásny T., Zimová S., Merganičová K., Štěpánek P., Modlinger R., Turčáni M. (2021): Devastating outbreak of bark beetles in the Czech Republic: Drivers, impacts, and management implications. Forest Ecology and Management, 490: 119075. Go to original source...
  61. Holuša J., Foit J., Knížek M., Schovánková J., Lukášová K., Vanická H., Trombík J., Kula E. (2019): The bark beetles Orthotomicus laricis and Orthotomicus longicollis are not pests in Central Europe: A case study from the Czech Republic. Bulletin of Insectology, 72: 253-260.
  62. Horgan T., Keane M., McCarthy R., Lally M., Thompson D., O'Carroll J. (2003): A Guide to Forest Tree Species Selection and Silviculture in Ireland. Dublin, National Council for Forest Research and Development (COFORD): 255.
  63. Jaehne S.C., Dohrenbusch A. (1997): Ein Verfahren zur Beurteilung der Bestandesdiversität. Forstwissenschaftliches Centralblatt, 116: 333-345. (in German) Go to original source...
  64. Jansson M., Fors H., Lindgren T., Wiström B. (2013): Perceived personal safety in relation to urban woodland vegetation - A review. Urban Forestry & Urban Greening, 12: 127-133. Go to original source...
  65. Johnson O., More D. (2006): Collins Tree Guide. London, Collins: 464.
  66. Knibbe B. (2007): PAST4: Personal Analysis System for Treering Research, Version 4.2. Vienna, SCIEM: 161.
  67. Knížek M., Liška J., Véle A., Zahradník P., Lubojacký J. (2021): Ochrana borovice lesní (Pinus sylvestris L.) před podkorním a dřevokazným hmyzem. Certifikovaná metodika, Lesnický průvodce 9/2021. Strnady, Výzkumný ústav lesního hospodářství a myslivosti: 120. (in Czech)
  68. Komonen A., Schroeder L.M., Weslien J. (2011): Ips typographus population development after a severe storm in a nature reserve in southern Sweden. Journal of Applied Entomology, 135: 132-141. Go to original source...
  69. Kovalevskii S.B., Krol A., Myroniuk V., Kovalevskyi S.S., Vysotska N., Khromulyak O., Yurchenko V. (2022): Growth of Scots pine (Pinus sylvestris L.) stands on soils with close bedding of crystalline parent rocks in Central Polissya, Ukraine. Central European Forestry Journal, 68: 72-77. Go to original source...
  70. Kraft G. (1884): Beiträgezur zur lehre von den durchforstungen, schlagstellungen und lichtungshieben. Hannover, Klindworth: 147. (in German)
  71. Krakau U.K. Liesebach M., Aronen T., Lelu-Walter M.A., Schneck V. (2013): Scots Pine (Pinus Sylvestris L.). Managing forest ecosystems. In: Pâques L.E. (ed.): Forest Tree Breeding in Europe: Current State-of-the-Art and Perspectives. Dordrecht, Springer: 267-323. Go to original source...
  72. Ledermann T., Neumann M. (2005): Ergebnisse vorläufiger Untersuchungen zur Erstellung von Biomassefunktionen aus Daten alter Dauerversuchsflächen. In: Deutscher Verband Forstlicher Forschungsanstalten, Sektion Ertragskunde, Beiträge zur Jahrestagung 2005, Freising, May 9-11, 2005: 143-152. (in German)
  73. Leštianska A., Fleischer Jr P., Merganičová K., Fleischer Sr P., Nalevanková P., Střelcová K. (2023): Effect of provenance and environmental factors on tree growth and tree water status of Norway spruce. Forests, 14: 156. Go to original source...
  74. Leuschner C., Ellenberg H. (2017): Ecology of Central European Forests: Vegetation Ecology of Central Europe, Volume 1. Berlin, Springer: 972. Go to original source...
  75. Lindner M., Maroschek M., Netherer S., Kremer A., Barbati A., Garciagonzalo J., Seidl R., Delzon S., Corona P., Kolström M., Marchetti M., Lexer M. (2010): Climate change impacts, adaptive capacity, and vulnerability of European forest ecosystems. Forest Ecology and Management, 259: 698-709. Go to original source...
  76. Liška J., Knížek M., Véle A. (2021): Evaluation of insect pest occurrence in areas of calamitous mortality of Scots pine. Central European Forestry Journal, 67: 85-90. Go to original source...
  77. Lubojacký J., Knížek M. (2021): Podkorní hmyz. In: Knížek M., Liška J. (eds): Výskyt lesních škodlivých činitelů v roce 2020 a jejich očekávaný stav v roce 2021. Zpravodaj ochrany lesa - Supplementum 2021. Strnady, Výzkumný ústav lesního hospodářství a myslivosti: 22-36. (in Czech)
  78. Matías L., Jump A.S. (2012): Interactions between growth, demography and biotic interactions in determining species range limits in a warming world: The case of Pinus sylvestris. Forest Ecology and Management, 282: 10-22. Go to original source...
  79. Mikeska M., Vacek S., Prausová R., Simon J., Minx T., Podrázský V., Malík V., Kobliha J., Anděl P., Matějka K. (2008): Typologické vymezení, struktura a management přirozených borů a borových doubrav v ČR. Kostelec nad Černými lesy, Lesnická práce: 450. (in Czech)
  80. Mikulenka P., Prokůpková A., Vacek Z., Vacek S., Bulušek D., Simon J., Šimůnek V., Hájek V. (2020): Effect of climate and air pollution on radial growth of mixed forests: Abies alba Mill. vs. Picea abies (L.) Karst. Central European Forestry Journal, 66: 23-36. Go to original source...
  81. Nemani R.R., Keeling C.D., Hashimoto H., Jolly W.M., Piper S.C., Tucker C.J., Myneni R.B., Running S.W. (2003): Climate-driven increases in global terrestrial net primary production from 1982 to 1999. Science, 300: 1560-1563. Go to original source... Go to PubMed...
  82. Novák J., Dušek D., Kacálek D., Slodičák M., Souček J. (2017): Pěstební postupy pro borové porosity 1. a 2. lesního vegetačního stupně. Strnady, Lesnický průvodce: 28. (in Czech)
  83. O'Reilly-Wapstra J.M., Moore B.D., Brewer M., Beaton J., Sim D., Wiggins N.L., Iason G.R. (2014): Pinus sylvestris sapling growth and recovery from mammalian browsing. Forest Ecology and Management, 325: 18-25. Go to original source...
  84. OECD (2006): Safety Assessment of Transgenic Organisms: OECD Consensus Documents, Vol. 2. Paris, OECD Publishing: 448. Go to original source...
  85. Petráš R., Pajtík J. (1991): Sústava česko-slovenských objemových tabuliek drevín. Lesnícky časopis, 37: 49-56. (in Slovak)
  86. Petráš R., Košút M., Oszlányi J. (1985): Listová biomasa stromov smreka, borovice a buka. Lesnícky časopis, 31: 121-135. (in Slovak)
  87. Plíva K. (2000): Trvale udržitelné obhospodařování lesů podle souborů lesních typů. Brandýs nad Labem, ÚHÚL: 200. (in Czech)
  88. Podrázský V., Remeš J. (2007): Změny kvality a množství nadložního humusu při přirozeném zmlazení bukových porostů na území Školního lesního podniku Kostelec nad Černými lesy. Zprávy lesnického výzkumu, 52: 118-122. (in Czech)
  89. Podrázský V., Remeš J. (2010): Effects of the species composition change on the humus form state in the forest stands on the territory of the University Training Forest at Kostelec nad Černými lesy. Zprávy lesnického výzkumu, 55: 71-77. Go to original source...
  90. Podrázský V., Remeš J., Kratochvíl J. (2005): Vývoj půdního chemizmu ve smrkových lesních ekosystémech na území ŠLP Kostelec nad Černými lesy. Zprávy lesnického výzkumu, 50: 198-201. (in Czech)
  91. Poleno Z., Vacek S., Podrázský V., Remeš J., Mikeska M., Kobliha J., Bílek L. (2007): Pěstování lesů II. Teoretická východiska pěstování lesů. Kostelec nad Černými lesy, Lesnická práce: 464. (in Czech)
  92. Poleno Z., Vacek S., Podrázský V., Remeš J., Štefančík I., Mikeska M., Kobliha J., Kupka I., Malík V., Turčáni M., Dvořák J., Zatloukal V., Bílek L., Baláš M., Simon J. (2009): Pěstování lesů III. Praktické postupy pěstování lesů. Kostelec nad Černými lesy, Lesnická práce: 952. (in Czech)
  93. Praciak A., Pasiecznik N., Sheil D., van Heist M., Sassen M., Sousa Correia C., Dixon C., Fyson G.E., Rushforth K., Teeling C. (2013): The CABI Encyclopedia of Forest Trees. Wallingford, CABI: 536.
  94. Pretzsch H. (2006): Wissen nutzbar machen für das Management von Waldökosystemen. Allgemeine Forst Zeitschrift für Waldwirtschaft und Umweltvorsorge, 61: 1158-1159. (in German)
  95. Pretzsch H., Biber P. (2016): Tree species mixing can increase maximum stand density. Canadian Journal of Forest Research, 46: 1179-1193. Go to original source...
  96. Pretzsch H., Schütze G. (2016): Effect of tree species mixing on the size structure, density, and yield of forest stands. European Journal of Forest Research, 135: 1-22. Go to original source...
  97. Průša E. (2001): Pěstování lesů na typologických základech. Kostelec nad Černými lesy, Lesnická práce: 594. (in Czech)
  98. Putalová T., Vacek Z., Vacek S., Štefančík I., Bulušek D., Král J. (2019): Tree-ring widths as an indicator of air pollution stress and climate conditions in different Norway spruce forest stands in the Krkonoše Mts. Central European Forestry Journal, 65: 21-33. Go to original source...
  99. Reddy A.R., Chaitanya K.V., Vivekanandan M. (2004): Drought-induced responses of photosynthesis and antioxidant metabolism in higher plants. Journal of Plant Physiology, 161: 1189-1202. Go to original source... Go to PubMed...
  100. Réh J. (1978): Technika pestovania lesa vo výbernej sústave hospodárenia. In: Vyskot M. et al.: Pěstění lesů. Praha, SZN: 360-412. (in Slovak)
  101. Reich P.B., Oleksyn J. (2008): Climate warming will reduce growth and survival of Scots pine except in the far north. Ecology Letters, 11: 588-597. Go to original source... Go to PubMed...
  102. Reineke L.H. (1933): Perfecting a stand density index for even-aged forests. Journal of Agricultural Research, 46: 627-638.
  103. Remeš J. (2006): Transformation of even-aged spruce stands at the School Forest Enterprise Kostelec nad Černými lesy: Structure and final cutting of mature stand. Journal of Forest Science, 52: 158-171. Go to original source...
  104. Remeš J., Kozel J. (2006): Structure, growth and increment of the stands in the course of stand transformation in the Klokočná Forest Range. Journal of Forest Science, 52: 573-546. Go to original source...
  105. Remeš J., Bílek L., Novák J., Vacek Z., Vacek S., Putalová T., Koubek L. (2015): Diameter increment of beech in relation to social position of trees, climate characteristics and thinning intensity. Journal of Forensic Sciences, 61: 456-464. Go to original source...
  106. Remeš J., Pulkrab K., Bílek L., Podrázský V. (2020): Economic and production effect of tree species change as a result of adaptation to climate change. Forests, 11: 431. Go to original source...
  107. Ruiz-Peinado R., Pretzsch H., Löf M., Heym M., Bielak K., Aldea J., Barbieto I., Brazaitis G., Drössler L., Godvod K. et al. (2021): Mixing effects on Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.) productivity along a climatic gradient across Europe. Forest Ecology and Management, 482: 118834. Go to original source...
  108. Rybníček M., Čermák P., Žid T., Kolář T. (2010): Radial growth and health condition of Norway spruce (Picea abies (L.) Karst.) stands in relation to climate (Silesian Beskids, Czech Republic). Geochronometria, 36: 9-16. Go to original source...
  109. Saniga M., Szanyi O. (1998): Modely výberkových lesov vo vybraných lesných typoch a geografických celkoch Slovenska. Vedecké štúdie TU vo Zvolene, 4/1998/A. Zvolen, Technická univerzita Zvolen: 48. (in Slovak)
  110. Schweingruber F.H. (1996): Tree Rings and Environment Dendroecology. Birmensdorf, Swiss Federal Institute for Forest, Snow and Landscape Research: 609.
  111. Seidl R., Rammer W., Lexer M. (2011): Climate change vulnerability of sustainable forest management in the Eastern Alps. Climatic Change, 106: 225-254. Go to original source...
  112. Sevik H., Topacoglu O. (2015): Variation and inheritance pattern in cone and seed characteristics of Scots pine (Pinus sylvestris L.) for evaluation of genetic diversity. Journal of Environmental Biology, 36: 1125.
  113. Sidor C.G., Popa I., Vlad R., Cherubini P. (2015): Different tree-ring responses of Norway spruce to air temperature across an altitudinal gradient in the Eastern Carpathians (Romania). Trees, 29: 985-997. Go to original source...
  114. Šimůnek V., Vacek Z., Vacek S. (2020): Solar cycles in salvage logging: National data from the Czech Republic confirm significant correlation. Forests, 11: 973. Go to original source...
  115. Souček J., Tesař V. (2008): Metodika přestavby smrkových monokultur na stanovištích přirozených smíšených porostů: Certifikovaná metodika. Lesnický průvodce, 4: 37. (in Czech)
  116. Souček J., Špulák O., Dušek D. (2018): Metodika přeměny a přestavby borových monokultur na stanovištích přirozených smíšených porostů: Certifikovaná metodika. Lesnický průvodce, 15: 35. (in Czech)
  117. Soulé P.T., Knapp P.A. (2006): Radial growth rate increases in naturally occurring ponderosa pine trees: A late-20th century CO2 fertilization effect? New Phytologist, 171: 379-390. Go to original source... Go to PubMed...
  118. Spathelf P., Ammer C. (2015): Forest management of Scots pine (Pinus sylvestris L.) in northern Germany - A brief review of the history and current trends. Forstarchiv, 86: 59-66.
  119. Spiecker H., Hansen J., Klimo E., Skovsgaard J.P., Streba H., Teuffel K.V. (2004): Norway spruce conversion - Options and consequences. European Forest Institute Research Report 18. Leiden, Brill: 269. Go to original source...
  120. Stanturf J.A., Palik B.J., Dumroese R K. (2014): Contemporary forest restoration: A review emphasizing function. Forest Ecology and Management, 331: 292-323. Go to original source...
  121. Švec O., Bílek L., Remeš J., Vacek Z. (2015): Analysis of operational approach during forest transformation in Klokočná Range, Central Bohemia. Journal of Forest Science, 61: 148-155. Go to original source...
  122. Sydorenko S., Voron V., Koval I., Sydorenko S., Rumiantsev M., Hurzhii R. (2021): Postfire tree mortality and fire resistance patterns in pine forests of Ukraine. Central European Forestry Journal, 67: 21-29. Go to original source...
  123. Taeger S., Zang C., Liesebach M., Schneck V., Menzel A. (2013): Impact of climate and drought events on the growth of Scots pine (Pinus sylvestris L.) provenances. Forest Ecology and Management, 307: 30-42. Go to original source...
  124. Tangwa E., Pechanec V., Brus J., Vyvlecka P. (2022): Spatial shifts in species richness in response to climate and environmental change: An adaption of the EUROMOVE model in the Czech Republic. Diversity, 14: 235. Go to original source...
  125. Teuffel K., Baumgarten M., Hanewinkel M., Sauter U.H., Spiecker H., von Wilpert K. (2005): Waldumbau: Für eine zukunftsorientierte Waldwirtschaft. Berlin, Springer: 422. (in German) Go to original source...
  126. Usoltsev V.A., Merganičová K., Konôpka B., Tsepordey I.S. (2022): The principle of space-for-time substitution in predicting Picea spp. biomass change under climate shifts. Central European Forestry Journal, 68: 174-189. Go to original source...
  127. Vacek S., Simon J., Remeš J., Podrázský V., Minx T., Mikeska M., Malík V., Jankopvský L., Turčáni M., Jakuš R., Schwarz O., Kozel J., Valenta M., Lička L., Hlásný T., Zúbrík M., Krejčí F., Třešňák J., Hofmeister Š. (2007): Obhospodařování bohatě strukturovaných a přírodě blízkých lesů. Kostelec nad Černými lesy, Lesnická práce: 447. (in Czech)
  128. Vacek S., Vacek Z., Bílek L., Simon J., Remeš J., Hůnová I. (2016): Structure, regeneration and growth of Scots pine (Pinus sylvestris L.) stands with respect to changing climate and environmental pollution. Silva Fennica, 50: 1564. Go to original source...
  129. Vacek Z., Vacek S., Eşen D., Yildiz O., Král J., Gallo J. (2020): Effect of invasive Rhododendron ponticum L. on natural regeneration and structure of Fagus orientalis Lipsky forests in the Black Sea region. Forests, 11: 603. Go to original source...
  130. Vacek Z., Linda R., Cukor J., Vacek S., Šimůnek V., Gallo J., Vančura K. (2021a): Scots pine (Pinus sylvestris L.), the suitable pioneer species for afforestation of reclamation sites? Forest Ecology and Management, 485: 118951. Go to original source...
  131. Vacek Z., Prokůpková A., Vacek S., Bulušek D., Šimůnek V., Hájek V., Králíček I. (2021b): Mixed vs. monospecific mountain forests in response to climate change: Structural and growth perspectives of Norway spruce and European beech. Forest Ecology and Management, 488: 119019. Go to original source...
  132. Vacek Z., Vacek S., Cukor J. (2023): European forests under global climate change: Review of tree growth processes, crises and management strategies. Journal of Environmental Management, 332: 117353. Go to original source... Go to PubMed...
  133. van der Maaten-Theunissen M., van der Maaten E., Kahle H.-P. (2013): Drought sensitivity of Norway spruce is higher than that of silver fir along an altitudinal gradient in southwestern Germany. Annals of Forest Science, 70: 185-193. Go to original source...
  134. Viewegh J., Kusbach A., Mikeska M. (2003): Czech forest ecosystem classification. Journal of Forest Science, 49: 74-82. Go to original source...
  135. Vítámvás J., Bílek L., Ulbrichová I., Bažant V., Dreslerová J., Vacek Z. (2019): Vzcházení, přežívání a kořenový systém semenáčků borovice lesní (Pinus sylvestris L.) při různých intenzitách slunečního záření a závlahy. Zprávy lesnického výzkumu, 64: 102-110. (in Czech)
  136. Vorčák J., Merganič J., Saniga M. (2006): Structural diversity change and regeneration processes of the Norway spruce natural forest in Babia hora NNR in relation to altitude. Journal of Forensic Sciences, 52: 399-409. Go to original source...
  137. Yamaguchi D.K. (1991): A simple method for cross-dating increment cores from living trees. Canadian Journal of Forest Research, 21: 414-416. Go to original source...

This is an open access article distributed under the terms of the Creative Commons Attribution-NonCommercial 4.0 International (CC BY NC 4.0), which permits non-comercial use, distribution, and reproduction in any medium, provided the original publication is properly cited. No use, distribution or reproduction is permitted which does not comply with these terms.