J. For. Sci., 2005, 51(5):203-212 | DOI: 10.17221/4591-JFS

Effect of the position in a stem on the length of tracheids in spruce (Picea abies [L.] Karst.) with the occurrence of reaction wood

V. Gryc, P. Horáček
Faculty of Forestry and Wood Technology, Mendel University of Agriculture and Forestry Brno, Brno, Czech Republic

The paper was aimed at the determination of variability of tracheid dimensions in spruce wood in relation to the position in a spruce stem. Significant changes in dimensions were found in early-wood and late-wood tracheids along the stem length and radius. There were statistically significant differences (variability along the radius) between particular annual rings. The height of 22 m showed statistically significant differences as compared with other heights (variability along the height). Differences between the length of early-wood and late-wood tracheids were not corroborated in zones CW, OW and SWL. Only in the SWP zone, statistically significant differences were found. Data sets (early-wood and late-wood tracheids) from the CW zone showed statistically significant differences as compared with other zones. On the basis of the results obtained, 3D models were created (for CW, OW, SWL and SWP zones; models for an early-wood and late-wood tracheid) describing changes in tracheid dimensions in spruce in relation to the position in a stem. In the models, the length of tracheids decreases with the height of a stem and on the other hand, with an increasing distance from the stem pith the length of tracheids increases. The importance of the paper consists in the enlargement of findings on the structure of spruce wood. In addition to this, the paper can contribute to the partial explanation of the different behaviour of physical and mechanical properties of wood in particular parts of the spruce stem.

Keywords: spruce; tracheid; length of tracheids; compression wood

Published: May 31, 2005  Show citation

ACS AIP APA ASA Harvard Chicago Chicago Notes IEEE ISO690 MLA NLM Turabian Vancouver
Gryc V, Horáček P. Effect of the position in a stem on the length of tracheids in spruce (Picea abies [L.] Karst.) with the occurrence of reaction wood. J. For. Sci. 2005;51(5):203-212. doi: 10.17221/4591-JFS.
Download citation

References

  1. BALABÁN K., 1955. Nauka o dřevě. Praha, SZN: 216.
  2. BUTTERFIELD B.G., MEYLAN B.A., 1980. Tree-Dimensional Structure of Wood. London, New York, Chapmann and Hall: 103. Go to original source...
  3. CASPERSON G., 1962. Über die Bildung der Zellwand beim Reaktionsholz. Holztechnologie, 3: 217-223.
  4. DOUDA V., 1948. Studie o významu reakčního dřeva. Praha, Sborník ČZA: 48.
  5. GÖTSCHE H., KÜHN H., 1988. Bildung und Eigenschaften des Holzes von Fichten (Picea abies /L./ Karst.) aus Waldschadensgebiet, Mitteilungen der Bundesforschungsamt für Forst und Holzwirtschaft. Hamburg, Kommissionverlag - Buchhandlung Max Wiedebusch: 171.
  6. GRYC V., HORÁČEK P., 2003. Vliv polohy ve kmeni na morfologii tracheid u smrku (Picea abies /L./ Karst.). In: Konference doktorandů MENDELNET 2003. Brno, MZLU, LDF: 53-58.
  7. CHU L., 1972. Comparison of normal wood and first-year compression wood in longleaf pine trees. Mississippi State University: 72.
  8. KNIGGE W., WENZEL B., 1982. Über die variabilität der Faserlänge innerhalb eines Stammes von Sequoiadendron giganteum (Lindl.) Buchholz. Forstarchiv, 3: 94-99.
  9. KNIGGE W., 1958. Das Phänomen der Reaktionsholzbildung und seine Bedeutung für die Holwervendung. Forstarchiv, 29: 4-10.
  10. KOLLMAN F., 1951. Technologie des Holzes und der Holzwerkstoffe. Berlin, Springer Verlag: 1048.
  11. KUČERA L.J., BOSSHARD H.H., 1989. Holzeigenschaften geschdigter Fichten. Basel, Berhuser Verlag: 182. Go to original source...
  12. MATOVIČ A., GANDELOVÁ L., 1980. Charakteristika dřeva smrku. In: Sborník z mezinárodní vědecké konference. Brno, V©Z, LF: 85-90.
  13. ONAKA F., 1949. Studies on compression and tension wood. Kyoto University, Mokuzai Kenkyo Wood Research Institute: 88.
  14. PANSHIN A.J., DE ZEEUW C., 1980. Textbook of Wood Technology. New York, McGraw-Hill, Inc.: 722.
  15. PLOMION CH., LEPROVOST G., STOKES A., 2001. Wood formation in trees. Plant Physiology, 127: 1513-1523. Go to original source...
  16. SANIO K., 1873. Anatomie der gemeinen Kiefer (Pinus sylvestris L.). Jahrbücher für wissenschaftliche Botanik, 9: 50-126.
  17. SARÉN M., SERIMAA R., ANDERSSON S., PAAKKARI T., SARANPÄÄ P., PESONEN E., 2001. Structural variation of tracheids in Norway spruce (Picea abies [L.] Karst.). Journal of Structural Biology, 136: 101-109. Go to original source... Go to PubMed...
  18. SACHSSE H., 1982. Holzeigenschaften der Weißfichte Picea glauca (Moench) Voss aus einem westdeutschen Versuchsanbau. Forstarchiv, 53: 21-24.
  19. SETH M.K., AGRAWAL H.O., 1984. Variation in tracheid length in blue pine (Pinus wallichiana A.B. Jackson). Holzforschung, 38: 1-6. Go to original source...
  20. SETH M.K., JAIN K.K., 1977. Relationship between percentage of compression wood and tracheid length in blue pine (Pinus wallichiana A.B. Jackson). Holzforschung, 31: 80-83. Go to original source...
  21. SEELING U., 1999. Einfluss von Richtgewebe (Druckholz) auf Festigkeit und Elastizitt des Druckholes. Holz als Roh- und Werkstoff, 57: 81-91. Go to original source...
  22. SHELBOURNE C., RITCHIE K., 1968. Relationship between degree of compression wood development and specific gravity and tracheid characteristic in loblolly pine (Pinus taeda L.). Holzforschung, 22: 185-190. Go to original source...
  23. SCHWEINGRUBER F.H., 1993. Jahrringe und Umwelt - Dendroökolgie. Bimensdorf, Eidgenössiche Forschungsanstalt für Wald, Schnee und Landschaft: 474.
  24. SISKO M., PFÄFFLI I., 1995. Fiber Atlas: Identification of Papermaking Fibers. Berlin, Springer Verlag: 400.
  25. ©KRIPEŇ J., RIASOVÁ T., 1958. Vlastnosti dreva borovice obyčajnej na Slovensku. Drevársky výskum, 3: 27-48.
  26. TIMELL T.E., 1986. Compresion Wood in Gymnosperms, Volume 1. Bibliography, Historical Background, Determination, Structure, Chemistry, Topochemistry, Physical Properties, Origin and Formation of Compression Wood. Berlin, Springer Verlag: 705.
  27. TRENDELENBURG R., 1939. Das Holz als Rohstoff. München, Berlin, Lehmanns Verlag: 435.
  28. TRENDELENBURG R., 1932. Über die Eigenchaften des Rotoder Druckholz der Nadelhölzer. Allgemeine Forst- und Jagdzeitung, 108: 1-14.
  29. WAGENFÜHR R., 1989. Anatomie des Holzes unter besondere Berücksichtigung der Holztechnik. Leipzig, VEB Fachbuchverlag: 334.
  30. WAGENFÜHR R., 1999. Anatomie des Holzes, Strukturanalytik - Identifizierung - Nomeklatur - Mikrotechnologie. Leipzig, DRW-Verlag: 188.
  31. YANG K.C., HAZENBERG G., 1994. Impact of spacing on tracheid length, relative density, and growth rate of juvenile wood and mature wood in Picea mariana. Canadian Forest Research, 24: 996-1007 Go to original source...

This is an open access article distributed under the terms of the Creative Commons Attribution-NonCommercial 4.0 International (CC BY NC 4.0), which permits non-comercial use, distribution, and reproduction in any medium, provided the original publication is properly cited. No use, distribution or reproduction is permitted which does not comply with these terms.